Karoo Lark
Staff, Students & Associates
Staff & Student Pages
Research Programmes



Online Public Access Catalogue & Reprints

The Niven Library's online public access catalogue is a searchable database listing all publications in the Library. Reprints can be obtained by contacting the Library Manager.

Marion and Prince Edward – Africa’s southern islands
Sinclair, I., Hockey, P., Ryan, P., Tarboton, W. 2011. Sasol Birds of Southern Africa IV. Struik Nature, Cape Town. 464 pp. ISBN: 9781770079250 (Also in Afrikaans)
Marion and Prince Edward – Africa’s southern islands
Terauds, A. Cooper, J. Chown, S.L. & Ryan, P. 2010. Marion & Prince Edward - Africa's Southern Islands. SUN Press, Stellenbosch. 176pp. ISBN:  978-1-920338-42-8.
Sinclair, I. & Ryan, P.G. 2009. Complete Photographic Field Guide: Birds of Southern Africa. Struik Nature, Cape Town. 432 pp. (Also in Afrikaans)
Sinclair, I. & Ryan, P.G. 2009. Complete Photographic Field Guide: Birds of Southern Africa. Struik Nature, Cape Town. 432 pp. 9781770073883 (Also in Afrikaans)
Staff, Students & Associates


Prof. Peter G. Ryan
BSc (Hons), MSc, PhD (Cape Town)

John Day Building: 2.06
Tel:  +27 (0)21 650 2966
Fax: +27 (0)21 650 3295

Email: peter.ryan@uct.ac.za

Activities and research interests

Academic Course Coordinator for the Institute's MSc Course in Conservation Biology, general ornithology, seabird-fishery interactions, evolutionary ecology (focusing on resolving closely-related species complexes), marine debris, solid waste management, the biology of oceanic islands.

Peter Ryan graduated from the University of Cape Town, researching the impacts of ingested plastic on seabirds (MSc) and the evolutionary ecology of buntings in the Tristan archipelago (PhD). After a post-doc at the University of California he was appointed academic Coordinator of the Masters Programme in Conservation Biology at the Fitztitute in 1993. Although he mostly works on seabirds and their conservation, he has wide-ranging interests in avian biology, and still maintains an interest in plastic pollution, especially in marine systems. He is the President of BirdLife South Africa and an associate editor of Antarctic Science and Bird Conservation International. He has been an honorary Conservation Officer at Tristan da Cunha since 1989, and is a member of Tristan’s Biodiversity Advisory Group. He has spent more than a year on Inaccessible Island, studying the island's endemic buntings as well as conducting surveys of threatened birds and conducting control programmes against alien plants. He is a keen birder and believes in the importance of promoting amateur involvement in ornithology. He has written several books about birds and their identification, and is scientific to African Birdlife, a popular magazine about birds produced by BirdLife South Africa.

Research programmes

Island conservation, Seabird conservation, Systematics and biogeography, Rarity & conservation of African birds, Co-operative breeding and sociality in birds

Current students


Dr Susan Cunningham
Dr Tom Flower
Dr Martha Nelson-Flower


Kate Carstens: Breeding dynamics, dispersal and fine-scale genetic structure in the endangered Southern Ground-Hornbill Bucorvus leadbeateri.  (Supervisors: Peter Ryan, Alan Kemp)

Alistair McInnes: At-sea behavioural responses of African Penguins in relation to small-scale variability in prey distribution: implications for Marine Protected Areas (Supervisors: Lorien Pichegru, Peter Ryan).

Lisa Nupen: Comparative conservation genetics and evolutionary history of threatened, endemic southern African seabirds in the Benguela Current Upwelling Ecosystem: Range-wide phylogeography, gene flow and population genetics based on nuclear and mitochondrial DNA (Supervisors: Peter Ryan, Rauri Bowie (UCLA, Berkeley), Jacqui Bishop (UCT, Zoology)).

Dominic Rollinson: Reducing seabird bycatch from the longline fishing industry (Supervisors: Peter Ryan, Ross Wanless)

Otto Whitehead: Foraging ecology and diving behaviour of crested penguins at Marion Island (Supervisors: Peter Ryan, Yan Ropert-Coudert).


Philna Botha: Testing the food-shortage hypothesis on the long-term population dynamics of the endangered Bank Cormorant (Supervisors: Timothée Cook, Richard Sherley, Les Underhill, Peter Ryan).

Davide Gaglio: Link between population dynamics, foraging behaviour and food abundance and distribution in the Swift Tern Thalasseus bergii (Supervisors: Timothée Cook, Richard Sherley, Peter Ryan).

Amanda Kyne: The allometry of feathers (Supervisor: Peter Ryan)

Corlia Meyer: Testing the influence of heat-stress on the breeding success of an endangered species: the Bank Cormorant (Supervisors:
Timothée Cook, Richard Sherley, Les Underhill, Peter Ryan).

Dane Paijmans: Changing breeding syndrome of African Black Oystercatchers (Supervisor: Peter Ryan).

John Pallett: Collisions of large birds, especially bustards, with power lines in Namibia: significance and possible solutions (Supervisors: Peter Ryan, Ann Scott).

Jenni Roberts: Tracking African Penguins in the non-breeding season (Supervisors: Peter Ryan, Ross Wanless, Lorien Pichegru, Christina Moseley)

Stefan Schoombie The breeding success and foraging ecology of Phoebetria spp. on Marion Island. (Supervisor: Peter Ryan).

Kim Stevens: The foraging ecology and breeding success of Grey-Headed albatross Thalassarche chrysostoma on Marion Island. (Supervisor: Peter Ryan)

Noelle Tubbs: Heat tolerance in the African Penguin (Spheniscus demersus) in the face of climate change (Supervisors: Lorien Pichegru, Peter Ryan, Jonathan Green).

Minke Witteveen: The breeding and foraging ecology of Kelp Gulls  (Supervisor: Peter Ryan).

Blair Zoghby: Conservation implications of patterns of habitat use by Southern Ground Hornbills (Supervisor: Peter Ryan)

Conservation Biology Masters

Salamatu Abdu: Spatial use of the landscape by Kalahari birds in relation to temperature and the location of water holes (Supervisors: Susie Cunningham, Peter Ryan)

Greg Campbell: Effects of temperature on gular fluttering and evaporative water loss in four sympatric cormorants in southern Africa (Supervisors: Tim Cook, Richard Sherley, Peter Ryan)

Binah Motlogelwa: Does climate niche predict bird population trends in South Africa? (Supervisors: Susie Cunningham, Peter Ryan)

Rukaya Johaadien: Aspects of the breeding biology of the Northern Rockhopper Penguin Eudyptes moseleyi.  (Supervisors: Peter Ryan, Antje Steinfurth)

Recent peer-reviewed publications

In press

Cohen, L.A., Pichegru, L., Grémillet, D., Coetzee, J., Upfold, L. and Ryan, P.G. (in press) A decade of changes in prey availability impact the foraging behaviour and fitness of Cape Gannets.  Marine Ecology Progress Series IP.

Gray, M.M., Wegmann, D., Haasl, R.J., White, M.A., Gabriel, S.I., Searle, J.B., Cuthbert, R.J., Ryan, P.G. & Payseur, B.A. (in press) Demographic history of a recent invasion of House Mice on the isolated island of Gough. Molecular Ecology IP.

Reid, T.A., Ronconi, R.A., Cuthbert, R.J. & Ryan, P.G. (in press) The summer foraging ranges of adult spectacled petrels Procellaria conspicillata. Antarctic Science IP. 

Ryan, P.G., Bourgeois, K., Dromzeé, S. and Dilley, B.J. (in press) The occurrence of two bill morphs of prions Pachyptila vittata on Gough Island. Polar Biology IP.

Ryan, P.G., Ortmann, H.E. & Herian, K. (in press) The impact of introduced scale insects on Phylica arborea and the implications for Nesospiza finches at Tristan da Cunha. Biological Conservation IP.

Schutgens, M., Shaw, J.M. & Ryan, P.G. (in press) Collision mortality of large birds on power lines: estimating scavenger and search bias in the Karoo, South Africa.  Ostrich IP.

Shaw, J.M., Dilley, B.J., Davies, D. and Ryan, P.G. (in press) Trapping and capture myopathy in Ludwig's Bustard. South African Journal of Wildlife Research IP.


Ryan, P.G. 2014.  Litter survey detects the South Atlantic 'garbage patch'. Marine Pollution Bulletin 79:220-224. doi: 10.1016/j.marpolbul.2013.12.010


Abad, F.X., Busquets, N., Sánchez, A., Ryan, P.G., Majó, N. & González-Solís, J. (2013). Serological and virological surveys of Influenza A virus in Antarctic and sub-Antarctic penguins. Antarctic Science 25:339-344.

Alström, P., Barnes, K.N., Olsson, U., Barker, F.K., Bloomer, P., Khan, A.A., Qureshi, M.A., Guillaumet, A., Crochet, P-A. & Ryan, P.G. (2013). Multilocus phylogeny of the avian family Alaudidae (larks) reveals 4 complex morphological evolution, non-monophyletic genera and hidden 5 species diversity. Molecular Phylogenetics and Evolution 69:1043-1056. 

Barquete, V., Strauss, V. & Ryan, P.G. (2013). Stable isotope turnover in blood and claws: A case study in captive African Penguins. Journal of Experimental Marine Biology and Ecology 448:121–127.

Commins, M.L., Ansorge, I. & Ryan, P.G. (2013) Multi-scale factors influencing seabird assemblages in the African sector of the Southern Ocean. Antarctic Science doi:10.1017./S0954102013000138.

Crawford, R.J.M., Randall, R.M., Whittington, P.A., Waller, L., Dyer, B.M., Allan, D.G., Fox, C., Martin, A.P., Upfold, L., Visagie, J., Bachoo, S., Bowker, M., Downs, C.T., Fox, R., Huisamen, J., Makhado, A.B., Oosthuizen, W.H., Ryan, P.G., Taylor, R.H. & Turpie, J.K. (2013). South Africa's coastal-breeding White-breasted Cormorants: population trends, breeding season and movements, and diet. African Journal of Marine Science 35: 473-490.

Cuthbert, R.J., Cooper, J. & Ryan, P.G. (2013). Population trends and breeding success of albatrosses and giant petrels at Gough Island in the face of at-sea and on-land threats. Antarctic Science doi.10.1017/S0954102013000424.

Cuthbert, R.J., Louw, H., Lurling, J., Parker, G., Rexer-Huber, K., Sommer, E., Visser, P. & Ryan, P.G. (2013). Low burrow occupancy and breeding success of burrowing petrels at Gough Island: a consequence of mouse predation. Bird Conservation International 23:113-124.

Dilley, B.J., Davies, D., Connan, M., Cooper, J., de Villiers, M., Swart, L., Vandenabeele, S., Ropert-Coudert, Y. & Ryan, P.G. (2013). Giant petrels as predators of albatross chicks. Polar Biology 36:761-766.

de Hierro, L., Moleón, M. & Ryan, P.G. (2013). Is carrying feathers a sexually selected trait in House Sparrows? Ethology 119:199-211.

Glass, J.P. & Ryan, P.G., (2013). Reduced Seabird night strikes and mortality in the Tristan rock lobster fishery. African Journal of Marine Science 35:589-592.

Jones, M.G.W. & Ryan, P.G. (2013). Effects of pre-laying attendance and body condition on long-term reproductive success in Wandering Albatrosses.  Emu http://dx.doi.org/10.1071/MU12054.

Le Roux, P.C., Ramaswiela, T., Kalwij, J.M., Shaw, J.D., Ryan, P.G., Treasure, A.M., McClelland, G.T.W., McGeoch, M.A. & Chown, S.L. (2013). Human activities, propagule pressure and alien plants in the sub-Antarctic: tests of generalities and evidence in support of management. Biological Conservation 161:18-27.

Little, R.M. & Ryan, P.G. (2013). Obituary: Philip Anthony Richard Hockey–'Education is what is left when what has been learnt has been forgotten' (8 March 1956–24 January 2013). Ostrich 84:v.

Little, R.M. & Ryan, P.G. (2013). Obituary: Philip Anthony Richard Hockey. African Zoology 48:197-198.

Little, R.M. & Ryan, P.G. (2013). Obituary: Phil Hockey: 8 March 1956 – 24 January 2013. South African Journal of Wildlife Research 43:87.

Reid, T.A. Ronconi, R.A., Cuthbert, R.J. & Ryan, P.G. (2013). Foraging range and habitat associations of non-breeding Tristan Albatrosses: overlap with fisheries and implications for conservation. Endangered Species Research 22:39-49.

Reid, T.A., Wanless, R.M., Hilton, G.M., Phillips, R.A. & Ryan, P.G. (2013). Foraging range and habitat associations of non-breeding Tristan albatrosses: overlap with fisheries and implications for conservation. Endangered Species Research 22:39-49.

Ryan, P.G. (2013). The effect of wind direction on flying fish counts. African Journal of Marine Science 35:585-587.

Ryan, P.G. (2013). Litter survey detects the South Atlantic ‘garbage patch’. Marine Pollution Bulletin 79:220-224. doi:10.1016/j.marpolbul.2013.12.010.

Ryan, P.G. (2013) Moult of flight feathers in darters (Anhingidae). Ardea 101(2):177-180.

Ryan, P.G. (2013) Medium-term changes in coastal bird communities inthe Western Cape, South Africa. Austral Ecology 38:251-259.

Ryan, P.G. (2013). A simple technique for counting marine debris at sea reveals steep litter gradients between the Straits of Malacca and the Bay of Bengal. Marine Pollution Bulletin 69:138-136.

Ryan, P.G. (2013). Medium-term changes in coastal bird communities in the Western Cape, South Africa. Austral Ecology 38:251-259.  IF 1.738

Ryan, P.G. (2013). Obituary: Phil Hockey (1956-2013). Ibis 155:698-700.

Ryan, P.G., Dyer, B.M., Martin, A.P., Ward, V.L., Whittington, P.A. & Williams, A.J. (2013). Supernormal clutches in southern African Kelp Gulls Larus dominicanus vetula. Ostrich 84:157-160.

Ryan, P.G., Klicka, L.B., Barker, K.F. & Burns, K.J. (2013). The origin of finches on Tristan da Cunha and Gough Island, central South Atlantic ocean. Molecular Phylogenetics and Evolution 69:299-305.

Ryan, P.G., Rose, B., Carter, M. & Clarke, R.H. (2013). A review of Jouanin’s Petrel records in the eastern Indian Ocean. Ostrich 84:161-164.

Tew-Kai, E., Benhamou, S., van der Lingen, C.D., Coetzee, J.C., Pichegru, L., Ryan, P.G. & Grémillet, D. (2013). Are Cape gannets dependent upon fishery waste? A multi-scale analysis using seabird GPS-tracking, hydro-acoustic surveys of pelagic fish and vessel monitoring systems. Journal of Applied Ecology 50:659-670.

Treasure, A.M., Moloney, C.L., Bester, M.N., McQuaid, C.D., Findlay, K.P., Best, P.B., Cowan, D.A., de Bruyn, P.J.N., Dorrington, R.A., Fagereng, A., Froneman, P.W., Grantham, G.H., Hunt, B.P.V., Meiklejohn, K.I., Pakhomov, E.A., Roychoudhury, A.N., Ryan, P.G., Smith, V.R., Chown, S.L. & Ansorge, I.J. (2013). South African research in the Southern Ocean: new opportunities and challenges. South African Journal of Science: doi:10.1590/sajs.2013/a009.


Chown, S.L., Lee, J.E., Hughes, K.A., Barnes, J., Barrett, P.J., Bergstrom, D.M., Convey, P., Cowan, D.A., Crosbie, K., Dyer, G., Frenot, Y., Grant, S.M., Herr, D., Kennicutt II, M.C., Lamers, M., Murray, A., Possingham, H.P., Reid, K., Riddle, M.J., Ryan, P.G., Sanson, L., Shaw, J.D., Sparrow, M.D., Summerhayes, C., Terauds, A. & Wall, D.H. (2012). Challenges to the Future Conservation of the Antarctic. Science 337:158-159.

Cook, T.R., Hamann, M., Pichegru, L., Bonadonna, F., Grémillet, D. & Ryan, P.G. (2012). GPS and time-depth loggers reveal underwater foraging plasticity in a flying diver, the Cape Cormorant. Marine Biology 159:373-387.

Hamann, M., Grémillet, D., Ryan, P.G., Bonadonna, F., van der Lingen, C.D. & Pichegru, L. (2012). A hard-knock life: the foraging ecology of Cape cormorants amidst shifting prey resources and industrial fishing pressure. African Journal of Marine Science 34:233-240.

Jansen van Rensburg, A., Bloomer, P., Ryan, P.G. & Bengt Hansson, B. (2012). Genetic variation at the major histocompatibility complex (MHCIIbeta) in Nesospiza buntings: Effects of genetic drift and shared ancestral polymorphism. BMC Evolutionary Biology 12:e143.

Jones, M.G.W., Techow, N.M.S.M. & Ryan, P.G. (2012). Dalliances and doubtful dads: what determines extra-pair paternity in socially monogamous wandering albatrosses? Behavioral Ecology and Sociobiology 66:1213-1224.

Moseley, C., Grémillet, D., Connan, M., Ryan, P.G., Mullers, R.H.E., der Lingen, C.D, Miller, T.W., Coetzee, J.C., Crawford, R.M., Sabarros, P., McQuaid, C.D. & Pichegru, L. (2012). Foraging ecology and ecophysiology of Cape gannets from colonies in contrasting feeding environments. Journal of Experimental Marine Biology and Ecology 422–423:29–38.

Parker, G.C., Rexer-Huber, K., Parker, K.A., Ryan, P.G., Techow, N.M.S.M. & Cuthbert, R.J. (2012). Morphometric and genetic sexing in the Gough Moorhen Gallinula comeri. Ringing and Migration 27: 20-25

Pichegru, L., Ryan, P.G., van Eeden, R., Reid, T., Grémillet, D. & Wanless, R., (2012). Industrial fishing, no-take zones and endangered penguins. Biological Conservation 156:117-125.

Rexer-Huber, K., Parker, G.C., Ryan, P.G. & Cuthbert, R.J. (2012). Ringing injuries in the Gough Moorhen Gallinula comeri. Ringing & Migration 27:53-56.

Ryan, P.G. (2012). Blue-breasted Kingfisher Halcyon malimbica feeding at sea. Malimbus 34:43-45.

Ryan, P.G. & Branch, G.M. (2012). The November 2011 irruption of buoy barnacles Dosima fascicularis in the Western Cape, South Africa. African Journal of Marine Science 34:157-162.


Cooper, J., Cuthbert, R.J., N.J.M. Gremmen, N.J.M., Ryan, P.G. & Shaw, J.D. 2011. Earth, fire and water: applying novel techniques to eradicate the invasive plant, procumbent pearlwort Sagina procumbens, on Gough Island, a World Heritage Site in the South Atlantic. In Veitch, C. R.; Clout, M. N. and Towns, D. R. (eds.). (2011). Proceedings of the Conference on Island invasives: eradication and management. IUCN, Gland, Switzerland. Pp. 162-165.

Crawford, R.J.M., Altwegg, R., Barham, B.J., Barham, P.J., Durant, J.M., Dyer, B.M., Geldenhuys, D., Makhado, A.B., Pichegru, L., Ryan, P.G.,Underhill, L.G., Upfold, L., Visagie, J.,Waller, L.J. & Whittington, P.A. (2011). Collapse of South Africa’s penguins in the early 21st century. African Journal of Marine Science 2011 33:139-156.

Crawford, R.J.M. & Ryan, P.G. (2011). Seabirds: sentinels of southern Africa’s seas. In Zietsman, L. (ed.) Observations on environmental change in South Africa. SUN Press, Stellenbosch. Pp. 247-257.

Cuthbert, R.J., Visser, P., Louw, H., Rexer-Huber, K., Parker, G., & Ryan, P.G. (2011). Preparations for the eradication of mice from Gough Island: results of bait acceptance trials above ground and around cave systems. In Veitch, C. R.; Clout, M. N. and Towns, D. R. (eds.). 2011. Proceedings of the Conference on Island invasives: eradication and management. IUCN, Gland, Switzerland. pp. 47-50.

Cuthbert, R.J., Visser, P., Louw. H. & Ryan, P.G. (2011). Palatability and efficacy of rodent baits for eradicating house mice (Mus musculus) from Gough Island, Tristan da Cunha. Wildlife Research 38:196-203.

Jenkins, A.R., Shaw, J.M., Smallie, J.J., Gibbons, B., Visagie, R. & Ryan, P.G. (2011). Estimating the impacts of power line collisions on Ludwig’s Bustards Neotis ludwigii. Bird Conservation International 21:303-310.

Pichegru, L., Ropert-Coudert, Y., Kato, A., Takahashi, A., Dyer, B.M. & Ryan, P.G. (2011). Diving patterns of female macaroni penguins breeding on Marion Island, South Africa. Polar Biology 34:945-954.

Robson, B., Glass, T., Glass, N., Glass, J., Green, J., Repetto, C., Rodgers, G., Ronconi, R.A., Ryan, P.G., Swain, G. & Cuthbert, R.J. (2011). Revised population estimate and trends for the Endangered Northern Rockhopper Penguin Eudyptes moseleyi at Tristan da Cunha. Bird Conservation International 21:454-459.

Ryan, P.G. & Ronconi, R.A. (2011). Continued increase in numbers of Spectacled Petrels Procellaria conspicillata. Antarctic Science 23:332-336.

Ryan, P.G., Glass, N. & Ronconi, R.A. (2011). The plants and birds of Stoltenhoff and Middle Islands, Tristan da Cunha. Polar Record 47:86-89.

Wright, K.L.B., Pichegru, L. & Ryan, P.G. (2011). Penguins are attracted to dimethyl sulphide at sea. Journal of Experimental Biology 214:2509-2511.


Cohen, C., Ryan, P.G., Claasen, M. & Ntoyinka, C. (2010). Birding Rwanda - in search of Red-collared Babbler Kupeornis rufocinctus, Albertine Rift endemics and more. Bulletin of the African Bird Club 17:229-237.

Grémillet, D., Mullers, R.H.E., Moseley, C., Pichegru, L., Coetzee, J.C., Sabarros, P.S., van der Lingen, C.D., Ryan, P.G., Kato, A. & Ropert-Coudert, Y. (2010). Seabirds, fisheries, and cameras. Frontiers in Ecology and the Environment 8: 401-402.

Jones, M.G.W. & Ryan, P.G. (2010). Evidence of mouse attacks on albatross chicks on sub-Antarctic Marion Island. Antarctic Science 22:39-42.

Mwema, M.M., de Ponte Machado, M. & Ryan, P.G. (2010). Breeding seabirds at Dassen Island, South Africa: chances of surviving great white pelican predation. Endangered Species Research 9:125-131.

Pichegru, L., Ryan, P.G., Crawford, R.J.M., van der Lingen, C.D. & Grémillet, D. (2010). Behavioural inertia places a top marine predator at risk from environmental change in the Benguela upwelling system. Marine Biology 157: 537-544.

Pichegru, L., Grémillet, D., Crawford, R.J.M. & Ryan, P.G. (2010). Marine no-take zone rapidly benefits endangered penguin. Biology Letters 6:498-501.

Ronconi, R.A., Ryan, P.G. & Ropert-Coudert, Y. (2010). Diving of Great Shearwaters (Puffinus gravis) in cold and warm water regions of the South Atlantic Ocean. Plos One 5:e15508.

Ryan, P.G. (2010). First record of Chilean Skua Catharacta chilensis for the African region. Bulletin of the African Bird Club 17:217-219.

Ryan, P.G., Nupen, L., Rose, B. & Suleiman, A.S. (2010).  Geographic variation in Socotra Sparrows Passer insularis. Bulletin of the British Ornithologists’ Club 130:75-82.

Ryan, P.G., Pichegru, L., Ropert-Coudert, Y., Grémillet, D. & Kato, A. (2010). On a wing and a prayer: the foraging ecology of breeding Cape cormorants. Journal of Zoology 280:25-32.

Ryan, P.G. & Ronconi, R.A. (2010). The Tristan Thrush Nesocichla eremita as seabird predator. Ardea 98:247-250.

Shaw, J.M., Jenkins, A.R., Smallie, J.J. & Ryan, P.G. (2010). Modelling power-line collision risk for the Blue Crane Anthropoides paradiseus in South Africa. Ibis 152:590-599.

Shaw, J.M., Jenkins, A.R., Ryan, P.G. & Smallie. (2010). A preliminary survey of avian mortality on power lines in the Overberg, South Africa. Ostrich 81:109-113.

Techow, N.M.S.M., O’Ryan, C., Phillips, R.A., Gales, R., Marin, M., Patterson-Fraser, D., Quintana, F., Ritz, M.S., Thompson, D.R., Wanless, R.M., Weimerskirch, H. & Ryan, P.G. (2010). Speciation and phylogeography of giant petrels Macronectes Molecular Phylogenetics and Evolution 54:472-487.

Wanless, R.M., Cooper, J., Slabber, M.J. & Ryan, P.G. (2010). Risk assessment of birds foraging terrestrially at Marion and Gough Islands to primary and secondary poisoning by rodenticides. Wildlife Research 37:524-530.

Wilson, J.W., Burle, M.-H., Cuthbert, R., Stirnemann, R.L. & Ryan, P.G. (2010). Breeding success of Northern Rockhopper Penguins (Eudyptes moseleyi) at Gough Island, South Atlantic Ocean. Emu 110:137-141.


Last modified: 2014/08/27
Copyright: Percy FitzPatrick Institute of African Ornithology 2014
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